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Whereas the pistillate inflorescences of its wild relatives are distichously arranged spikes of 5 to 12 caryopses encased in hard fruitcases, alternately stacked one on top of the other, that shatter upon maturity for natural seed dispersal, the many-rowed maize ear bears hundreds of exposed kernels that occur in pairs in cupules fused to form a rigid, nonshattering rachis (the cob). The entire structure is enveloped by a multiple of condensed leaf sheaths (the husks). Maize has been so radically modified by artificial selection that the plant is no longer capable of naturally reproducing itself.

The specimen labelled “possible F1 corn × teosinte closely resembles an inflorescence of Tripsacum × Z. diploperennis. From the collections at the Harvard University Herbaria. Guatemalan teosinte (Z. luxurians) and the two perennial species (Z. perennis and Z. diploperennis). 1 compares the two taxonomic classifications and aligns them with the geographic races of annual teosinte described by Wilkes (1967, 1972). The teosintes today extend from northern Mexico to southern Guatemala and northern Nicaragua.

THE ORIGIN OF MAIZE: EVIDENCE FOR TRIPSACUM ANCESTRY 41 urians) and the perennials (Z. diploperennis and Z. perennis). From these data, it can be inferred that Z. m. ssp. e. Balsas teosinte) is most closely related to maize. Doebley (1990) interprets this as evidence that maize is descended directly from Balsas teosinte. Comparative isozyme data for Tripsacum have not been published. Chloroplast (cp) DNA restriction analysis (Doebley et al. 1987; Doebley 1990) complemented results of isozyme studies with the exception that it placed Z.

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